FIELD TRIP TO THE HAILE QUARRIES

JANUARY 22, 2000

By Richard Hulbert and Roger Portell

Commercial limestone quarrying operations northwest of Newberry, Alachua County, Florida have been active for over 50 years. The Eocene bedrock is mined to a depth of about 70 feet, extending below the local water table, using a combination of blasting and draglines. These activities have exposed many important fossil localities over the decades, sites that would otherwise never have been discovered. These quarries and the fossil sites they contain are known in the scientific literature as the Haile localities or Haile complex. The small village of Haile no longer exists, but its name live on in this manner.

To keep track of the numerous Haile fossil sites, UF paleontologists systematically number each quarry. Each individual fossil site within a quarry was assigned a letter. Thus a combination of a number and a letter uniquely identifies each locality. Until recently, the numbers were reported using Roman numerals, although in the early 1990s the Florida Museum of Natural History (FLMNH) switched to Arabic numerals (so Haile XIVD became Haile 14D).

Continuing mining operations usually destroy the fossil localities at Haile. Therefore, many of the classic sites no long exist. The entire region is private land and fossil collecting is not allowed without permission of both the landowners and mine operators.

HISTORY

Vertebrate remains often accumulate in caves, sinkholes, and fissures. Bones and teeth are rapidly buried, protected from erosion, and the chances of mineralization by ground water are relatively good in such environments. While a relatively small percentage of the earthís sediments and sedimentary rocks are deposited in these kinds of environments, they have produced many important fossil sites around the world. Much of Floridaís surficial bedrock is limestone. Dissolution by acidic groundwater has produced thousands of solution features (caves, sinkholes, and chimneys). This process has been on-going ever since the state emerged above sea level in the Oligocene Epoch, about 30 million years ago (Ma).

The sediments that filled the sinkholes and fissures in northern Florida are rarely exposed by natural processes. Human excavations, either by mining or construction, are necessary to bring them to light. In the first part of this century, there were scattered small-scale phosphate mines in northern peninsular Florida, including the vicinity of Haile. Phosphatic sediments accumulated in sinkholes and channels eroded into the limestone at a time of high sea level during the Miocene Epoch (24 to 5 Ma). These were the sources of the first vertebrate fossils reported from the Newberry-Haile region (Simpson, 1930). Following WWII, small-scale phosphate mining became unprofitable, but limestone quarrying began in the region instead. Sinkholes, fissures, and caves were exposed at a more prolific rate. Scientific publications on vertebrate fossils from Haile began to regularly appear in the early 1950s. This early work was primarily by UF paleontologists Walter Auffenberg and Pierce Brodkorb, or one of their students. Since then, several generations of UF faculty, staff, and students have collected fossils at Haile, and there is no end in sight. As recently as this September, S. D. Emslie published a major review of Pliocene and Pleistocene birds from Florida. Fossils from several Haile sites figured prominently in his work. Additionally, there is on going research on ground sloths, tapirs, armadillos, turtles, mollusks, crabs, corals, and echinoderms from Haile.

GEOLOGY AND GEOCHRONOLOGY

The bedrock in the Haile region consists of marine limestone named the Ocala Group. The Ocala Group was deposited in a shallow, quiet environment and is abundantly fossiliferous. Most common is the disk-shaped, benthic foraminiferan Nummulites, which reaches an impressive size for a single-celled organism. The age of the Ocala Group is middle to late Eocene, about 40 to 35 Ma. This geologic age is based on correlation using macro invertebrates and microfossils with well-dated rocks of the middle and western Gulf Coastal Plain.

During times of emergence above sea level, sinkholes, caves, and fissures form in this portion of the Florida peninsula. Fissures are narrow, crevasse-like openings in the bedrock, as opposed to the more familiar round sinkholes. Most of the fossiliferous fissures and sinkholes in the Haile region (about 66%) are late Pleistocene in age, probably dating between 10 and 25 thousand years old. They are usually filled with rust red-colored sand. Their fossils are mostly terrestrial mammals, reptiles, and birds; aquatic forms are rare or absent. The second most common age for Haile vertebrate sites is late Miocene (about 16%), approximately 8 Ma. Geologically, these frequently contain abundant phosphatic gravel and sand. Marine and estuarine fossils are most common, including fish, sharks, dugongs, and the crocodile Gavialosuchus. Terrestrial species are also present, especially horses and camels.

Although not found as frequently as late Pleistocene or Miocene sites, some of the largest and most important Haile sites are Pliocene or early Pleistocene (2.5 to 1 Ma). Most sites of this age have abundant clay, or alternating layers of clay and sand, mixed the limestone rubble and boulders. The latter represent collapses of sinkhole or cave walls. In contrast to the late Pleistocene sites, there is a strong aquatic influence to these faunas. It is usually an exclusively fresh water component, unlike the primarily marine nature of the Miocene sites. Pliocene or early Pleistocene Haile sites have produced most of the articulated skeletons from the region. Vertebrates of all sizes have been found, ranging from small bats and frogs to large sloths and proboscideans.

INVERTEBRATE FOSSILS FROM HAILE

The Eocene limestone produces most of the fossil invertebrates from the Haile quarries. Compressed, internal molds of Late Pliocene land snails from Haile 7C are the lone exception. At Haile, invertebrates with shells made of the mineral aragonite are typically preserved as either internal or external molds. This includes most gastropods and bivalves. Many of the species preserved in this manner are poorly understood and are presently under investigation by Roger W. Portell, an invertebrate paleontologist at the FLMNH. Recently, the quarries at Haile produced the first Eocene muricid snails from Florida, Pterynotus rogersi, and an undescribed species of sea hare (marine gastropods that lack or have reduced shells) in the genus Philine.

Invertebrates with calcite shells are typically better preserved. There are over 20 species of echinoids (sea urchins, sea biscuits, and sand dollars). Among the most common are Oligopygus haldemani and Oligopygus wetherbyi (sea biscuits) and Neolagnum durhami and Weisbordella cubae (sand dollars). Perhaps the most spectacular invertebrate species commonly collected from the Haile quarries is the fossil stone crab Ocalina floridana-dozens of complete specimens have been found. The fossils of Ocalina are preserved a dark blue or brown color, and so are relatively easy to find in the light tan limestone.

SHARKS AND FISH

The Eocene limestone produces marine fish and sharks, most commonly shark teeth such as Carcharodon "auriculatus", ray mouth plates, and porcupine fish jaws. The grooved, round spine or rostrum of Cylindracanthus, a possible relative of the swordfish, has also been found. Among the Miocene fish, tarpon, drum, and gar are most abundant. Common large sharks of the time include Carcharodon, Isurus, Hemipristis, and Carcharhinus. The freshwater fish of the Pliocene and Pleistocene include catfish and sunfish.

AMPHIBIANS AND REPTILES

A wide variety of frogs, salamanders, lizards, snakes, turtles, and crocodiles have been collected from Haile. The following is just a sampling of some of the more notable occurrences. The late Miocene Haile 19A site produced such an abundance of the long-snouted crocodile Gavialosuchus americanus that the locality was informally known as the ìcroc hole.î The late Pliocene Haile 7C and 15A sites both produced many specimens of turtles, including snapping, softshell, and cooter/slider varieties. The ancestors of many of Floridaís modern turtles were first found and described on the basis of Haile specimens. Land tortoises have also been found at Miocene, Pliocene, and Pleistocene sites. The rare Eocene sea snake Pterosphenus has been collected in the Eocene limestone.

BIRDS

Fossil birds, commonly thought of as rare, are at times quite abundant in sinkhole/fissure deposits. Several of the Haile localities have produced important fossil bird faunas. The late Pleistocene Haile 11B site contained 71 species, the early Pleistocene Haile 16A 30 species, and the Pliocene Haile 7C 15 species. Many well preserved fossils of a large anhinga and a cormorant were collected at the late Miocene Haile 19A site. The largest known condor (Aizenogyps toomeyae), an undescribed species of golden eagle, and the first record of the ringed kingfisher (Ceryle torquata) from North America all came from Haile 7C. Haile 16A produced abundant fossils of turkey, bobwhite quail, rails, barn owls, and screech owls. Quail were also very abundant at Haile 11B, with other common birds including 10 different species of ducks, pied-billed grebe, king rail, burrowing owl, and sparrow hawk.

MAMMALS

The Ocala Group at Haile has produced a limited number of specimens of the extinct archaeocete whales, such as Zygorhiza. The late Miocene Haile sites have produced essentially the same mammalian fauna as the Love Site near Archer and the McGehee Farm Site south of Newberry. Isolated horse teeth belonging to the genera Cormohipparion, , Neohipparion, Nannippus, Calippus, and others have been found. Also present were giraffe-camels, llama-like camels, rhinos, tapirs, and the large dog Epicyon validus.

Pliocene and early Pleistocene sites (Haile 7C, 15A, 16A, and 21A) are rich in mammalian fossils. Among the common large mammals found at these sites are ground sloths, armadillos, pampatheres, tapirs, three- and one-toed horses, peccaries, and mastodonts. The oldest species of vampire bat, Desmodus archaeodaptes, has been found at Haile 16A. The carnivores of this interval include saber-toothed cats (Smilodon and Homotherium), wolves (Canis), and large bears (Arctodus). The late Pleistocene mammals of Haile include numerous small forms such as rodents, rabbits, bats, shrews, and moles. Among the larger mammals of this time, frequent finds include deer, bison, peccary, llama, horse, and mammoth.

PARTIAL HAILE BIBLIOGRAPHY

Berta, A. 1987. The sabrecat Smilodon gracilis from Florida and a discussion of its relationships (Smilodontinae, Felidae, Mammalia). Bulletin of the Florida State Museum, 31:1-63.
Campbell, K. E. 1976. An early Pleistocene avifauna from Haile XVA, Florida. Wilson Bulletin, 88:345-347.
Emslie, S. D. 1998. Avian community, climate, and sea-level changes in the Plio-Pleistocene of the Florida Peninsula. Ornithological Monographs, 50:1-113.
Hulbert, R. C. 1988. Cormohipparion and Hipparion (Mammalia, Perissodactyla, Equidae) from the late Neogene of Florida. Bulletin of the Florida State Museum, 33:229-338.
Ligon, J. D. 1965. A Pleistocene avifauna from Haile, Florida. Bulletin of the Florida State Museum, 10:127-158. Morgan, G. S., and R. C.
Hulbert. 1995. Overview of the geology and vertebrate biochronology of the Leisey Shell Pit local fauna, Hillsborough County, Florida. Bulletin of the Florida Museum of Natural History, 37:1-92.
Morgan, G. S., O. J. Linares, and C. E. Ray. 1988. New species of fossil vampire bats (Mammalia: Chiroptera: Desmodontidae) from Florida and Venezuela. Proceeding of the Biological Society of Washington, 101:912-928.
Portell, R.W., and E.H. Vokes. A new species of Pterynotus (Gastropoda: Muricidae) from the Eocene Ocala Limestone of Florida. Tulane Studies in Geology and Paleontology, 30(3):203-206.
Robertson, J. S. 1976. Latest Pliocene mammals from Haile XVA, Alachua County, Florida. Bulletin of the Florida State Museum, 20:111-186.
Simpson, G. G. 1930. Tertiary land mammals of Florida. Bulletin of the American Museum of Natural History, 49:149-211.
Weaver, W. G., and J. S. Robertson. 1967. A re-evaluation of fossil turtles of the Chrysemys scripta group. Tulane Studies in Geology, 5:53-66.
Webb, S. D. 1974. Pleistocene Mammals of Florida. University Presses of Florida, Gainesville.