FIELD TRIP TO THE HAILE QUARRIES
JANUARY 22, 2000
By Richard Hulbert and Roger Portell
Commercial limestone quarrying operations northwest of Newberry, Alachua
County, Florida have been active for over 50 years. The Eocene bedrock
is mined to a depth of about 70 feet, extending below the local water table,
using a combination of blasting and draglines. These activities have exposed
many important fossil localities over the decades, sites that would otherwise
never have been discovered. These quarries and the fossil sites they contain
are known in the scientific literature as the Haile localities or Haile
complex. The small village of Haile no longer exists, but its name live
on in this manner.
To keep track of the numerous Haile fossil sites, UF paleontologists
systematically number each quarry. Each individual fossil site within a
quarry was assigned a letter. Thus a combination of a number and a letter
uniquely identifies each locality. Until recently, the numbers were reported
using Roman numerals, although in the early 1990s the Florida Museum of
Natural History (FLMNH) switched to Arabic numerals (so Haile XIVD became
Haile 14D).
Continuing mining operations usually destroy the fossil localities at
Haile. Therefore, many of the classic sites no long exist. The entire
region is private land and fossil collecting is not allowed without permission
of both the landowners and mine operators.
HISTORY
Vertebrate remains often accumulate in caves, sinkholes, and fissures.
Bones and teeth are rapidly buried, protected from erosion, and the chances
of mineralization by ground water are relatively good in such environments.
While a relatively small percentage of the earthís sediments and
sedimentary rocks are deposited in these kinds of environments, they have
produced many important fossil sites around the world. Much of Floridaís
surficial bedrock is limestone. Dissolution by acidic groundwater has produced
thousands of solution features (caves, sinkholes, and chimneys). This process
has been on-going ever since the state emerged above sea level in the Oligocene
Epoch, about 30 million years ago (Ma).
The sediments that filled the sinkholes and fissures in northern Florida
are rarely exposed by natural processes. Human excavations, either by mining
or construction, are necessary to bring them to light. In the first part
of this century, there were scattered small-scale phosphate mines in northern
peninsular Florida, including the vicinity of Haile. Phosphatic sediments
accumulated in sinkholes and channels eroded into the limestone at a time
of high sea level during the Miocene Epoch (24 to 5 Ma). These were the
sources of the first vertebrate fossils reported from the Newberry-Haile
region (Simpson, 1930). Following WWII, small-scale phosphate mining became
unprofitable, but limestone quarrying began in the region instead. Sinkholes,
fissures, and caves were exposed at a more prolific rate. Scientific publications
on vertebrate fossils from Haile began to regularly appear in the early
1950s. This early work was primarily by UF paleontologists Walter Auffenberg
and Pierce Brodkorb, or one of their students. Since then, several generations
of UF faculty, staff, and students have collected fossils at Haile, and
there is no end in sight. As recently as this September, S. D. Emslie published
a major review of Pliocene and Pleistocene birds from Florida. Fossils
from several Haile sites figured prominently in his work. Additionally,
there is on going research on ground sloths, tapirs, armadillos, turtles,
mollusks, crabs, corals, and echinoderms from Haile.
GEOLOGY AND GEOCHRONOLOGY
The bedrock in the Haile region consists of marine limestone named the
Ocala Group. The Ocala Group was deposited in a shallow, quiet environment
and is abundantly fossiliferous. Most common is the disk-shaped, benthic
foraminiferan Nummulites, which reaches an impressive size
for a single-celled organism. The age of the Ocala Group is middle to late
Eocene, about 40 to 35 Ma. This geologic age is based on correlation using
macro invertebrates and microfossils with well-dated rocks of the middle
and western Gulf Coastal Plain.
During times of emergence above sea level, sinkholes, caves, and fissures
form in this portion of the Florida peninsula. Fissures are narrow, crevasse-like
openings in the bedrock, as opposed to the more familiar round sinkholes.
Most of the fossiliferous fissures and sinkholes in the Haile region (about
66%) are late Pleistocene in age, probably dating between 10 and 25 thousand
years old. They are usually filled with rust red-colored sand. Their fossils
are mostly terrestrial mammals, reptiles, and birds; aquatic forms are
rare or absent. The second most common age for Haile vertebrate sites is
late Miocene (about 16%), approximately 8 Ma. Geologically, these frequently
contain abundant phosphatic gravel and sand. Marine and estuarine fossils
are most common, including fish, sharks, dugongs, and the crocodile Gavialosuchus.
Terrestrial species are also present, especially horses and camels.
Although not found as frequently as late Pleistocene or Miocene sites,
some of the largest and most important Haile sites are Pliocene or early
Pleistocene (2.5 to 1 Ma). Most sites of this age have abundant clay, or
alternating layers of clay and sand, mixed the limestone rubble and boulders.
The latter represent collapses of sinkhole or cave walls. In contrast to
the late Pleistocene sites, there is a strong aquatic influence to these
faunas. It is usually an exclusively fresh water component, unlike the
primarily marine nature of the Miocene sites. Pliocene or early Pleistocene
Haile sites have produced most of the articulated skeletons from the region.
Vertebrates of all sizes have been found, ranging from small bats and frogs
to large sloths and proboscideans.
INVERTEBRATE FOSSILS FROM HAILE
The Eocene limestone produces most of the fossil invertebrates from the
Haile quarries. Compressed, internal molds of Late Pliocene land snails
from Haile 7C are the lone exception. At Haile, invertebrates with shells
made of the mineral aragonite are typically preserved as either internal
or external molds. This includes most gastropods and bivalves. Many of
the species preserved in this manner are poorly understood and are presently
under investigation by Roger W. Portell, an invertebrate paleontologist
at the FLMNH. Recently, the quarries at Haile produced the first Eocene
muricid snails from Florida, Pterynotus rogersi, and an undescribed
species of sea hare (marine gastropods that lack or have reduced shells)
in the genus Philine.
Invertebrates with calcite shells are typically better preserved. There
are over 20 species of echinoids (sea urchins, sea biscuits, and sand dollars).
Among the most common are Oligopygus haldemani and Oligopygus
wetherbyi (sea biscuits) and Neolagnum durhami and Weisbordella
cubae (sand dollars). Perhaps the most spectacular invertebrate species
commonly collected from the Haile quarries is the fossil stone crab Ocalina
floridana-dozens of complete specimens have been found. The fossils
of Ocalina are preserved a dark blue or brown color, and so are
relatively easy to find in the light tan limestone.
SHARKS AND FISH
The Eocene limestone produces marine fish and sharks, most commonly shark
teeth such as Carcharodon "auriculatus", ray mouth plates, and porcupine
fish jaws. The grooved, round spine or rostrum of Cylindracanthus,
a possible relative of the swordfish, has also been found. Among the Miocene
fish, tarpon, drum, and gar are most abundant. Common large sharks of the
time include Carcharodon, Isurus, Hemipristis, and
Carcharhinus. The freshwater fish of the Pliocene and Pleistocene
include catfish and sunfish.
AMPHIBIANS AND REPTILES
A wide variety of frogs, salamanders, lizards, snakes, turtles, and crocodiles
have been collected from Haile. The following is just a sampling of some
of the more notable occurrences. The late Miocene Haile 19A site produced
such an abundance of the long-snouted crocodile Gavialosuchus americanus
that the locality was informally known as the ìcroc hole.î
The late Pliocene Haile 7C and 15A sites both produced many specimens of
turtles, including snapping, softshell, and cooter/slider varieties. The
ancestors of many of Floridaís modern turtles were first found and
described on the basis of Haile specimens. Land tortoises have also been
found at Miocene, Pliocene, and Pleistocene sites. The rare Eocene sea
snake Pterosphenus has been collected in the Eocene limestone.
BIRDS
Fossil birds, commonly thought of as rare, are at times quite abundant
in sinkhole/fissure deposits. Several of the Haile localities have produced
important fossil bird faunas. The late Pleistocene Haile 11B site contained
71 species, the early Pleistocene Haile 16A 30 species, and the Pliocene
Haile 7C 15 species. Many well preserved fossils of a large anhinga and
a cormorant were collected at the late Miocene Haile 19A site. The largest
known condor (Aizenogyps toomeyae), an undescribed species of golden
eagle, and the first record of the ringed kingfisher (Ceryle torquata)
from North America all came from Haile 7C. Haile 16A produced abundant
fossils of turkey, bobwhite quail, rails, barn owls, and screech owls.
Quail were also very abundant at Haile 11B, with other common birds including
10 different species of ducks, pied-billed grebe, king rail, burrowing
owl, and sparrow hawk.
MAMMALS
The Ocala Group at Haile has produced a limited number of specimens of
the extinct archaeocete whales, such as Zygorhiza. The late Miocene
Haile sites have produced essentially the same mammalian fauna as the Love
Site near Archer and the McGehee Farm Site south of Newberry. Isolated
horse teeth belonging to the genera Cormohipparion, , Neohipparion,
Nannippus, Calippus, and others have been found. Also present
were giraffe-camels, llama-like camels, rhinos, tapirs, and the large dog
Epicyon validus.
Pliocene and early Pleistocene sites (Haile 7C, 15A, 16A, and 21A) are
rich in mammalian fossils. Among the common large mammals found at these
sites are ground sloths, armadillos, pampatheres, tapirs, three- and one-toed
horses, peccaries, and mastodonts. The oldest species of vampire bat, Desmodus
archaeodaptes, has been found at Haile 16A. The carnivores of this
interval include saber-toothed cats (Smilodon and Homotherium),
wolves (Canis), and large bears (Arctodus). The late Pleistocene
mammals of Haile include numerous small forms such as rodents, rabbits,
bats, shrews, and moles. Among the larger mammals of this time, frequent
finds include deer, bison, peccary, llama, horse, and mammoth.
PARTIAL HAILE BIBLIOGRAPHY
Berta, A. 1987. The sabrecat Smilodon gracilis from Florida and
a discussion of its relationships (Smilodontinae, Felidae, Mammalia). Bulletin
of the Florida State Museum, 31:1-63.
Campbell, K. E. 1976. An early Pleistocene avifauna from Haile XVA,
Florida. Wilson Bulletin, 88:345-347.
Emslie, S. D. 1998. Avian community, climate, and sea-level changes
in the Plio-Pleistocene of the Florida Peninsula. Ornithological Monographs,
50:1-113.
Hulbert, R. C. 1988. Cormohipparion and Hipparion (Mammalia,
Perissodactyla, Equidae) from the late Neogene of Florida. Bulletin of
the Florida State Museum, 33:229-338.
Ligon, J. D. 1965. A Pleistocene avifauna from Haile, Florida. Bulletin
of the Florida State Museum, 10:127-158. Morgan, G. S., and R. C.
Hulbert. 1995. Overview of the geology and vertebrate biochronology
of the Leisey Shell Pit local fauna, Hillsborough County, Florida. Bulletin
of the Florida Museum of Natural History, 37:1-92.
Morgan, G. S., O. J. Linares, and C. E. Ray. 1988. New species of fossil
vampire bats (Mammalia: Chiroptera: Desmodontidae) from Florida and Venezuela.
Proceeding of the Biological Society of Washington, 101:912-928.
Portell, R.W., and E.H. Vokes. A new species of Pterynotus (Gastropoda:
Muricidae) from the Eocene Ocala Limestone of Florida. Tulane Studies in
Geology and Paleontology, 30(3):203-206.
Robertson, J. S. 1976. Latest Pliocene mammals from Haile XVA, Alachua
County, Florida. Bulletin of the Florida State Museum, 20:111-186.
Simpson, G. G. 1930. Tertiary land mammals of Florida. Bulletin of
the American Museum of Natural History, 49:149-211.
Weaver, W. G., and J. S. Robertson. 1967. A re-evaluation of fossil
turtles of the Chrysemys scripta group. Tulane Studies in Geology,
5:53-66.
Webb, S. D. 1974. Pleistocene Mammals of Florida. University Presses
of Florida, Gainesville.